Evolution of Excretory System

 


Sponges:

The main opening of a sponge is used as an exit to repel waste.

Hydra/Polyp (cnidarians):

The main opening of cnidarians serves as mouth and anus.

Flatworm/Tapeworm (Planarians):

 

Because planarians live in fresh water, which tends to enter the body by osmosis, they have an excretory organ that largely rids the body of excess water.

The organ consists of a series of interconnecting canals, which run the length of body on each side.

The beating of cilia in flame cells (so named because the beating of cilia reminded some early investigators of flickering of a flame) keeps the water moving toward the excretory pores.

Roundworm:

 

Circulatory and excretory food digested in gut is not distributed by any specialised vascular system. Nutrients and waste are distributed in body cavity whose contents are regulated by one celled glands (in simple species) or excretory canal along each side of the body.

Molluscs

 

Clam (Bivalve):

 

There are two excretory kidneys in clam, which lie just below the heart and remove waste from the pericardial cavity for excretion into the mantle cavity.

The clam excretes ammonia, a poisonous substance that requires the concomitant excretion of water. Land dwelling animals tend to excrete a less toxic substance in a more concentrated form.

Snail (Gastropod):

 

There are 2 kidneys or nephridia in only the primitive gastropods such as archea gastropods while in advanced forms one kidney is small or lost.

The kidney plays different roles, depending upon the environment in which snail lives.

Usually the posterior chamber excretes uric acid and purines, while the anterior chamber has osmoregulatory function.

Snails living on land are uricotelic, this means that to preserve water they excrete almost solid uric acid.

Snails living in water excrete ammonia and are not uricotelic.

The renal aperture (urine opening) is situated in upper region of right mantle cavity. The urine produced by kidneys is expelled here.

Terrestrial gastropods reduce water loss by sealing the mantle cavity with an extended mantle collar.

Octopus (Cephalopod):

 

Excretory functions are carried out by nephridia (kidneys), tubular structures that collect fluid from the coelom and exchange salts and other substances with body tissues as fluid passes along the tubules for excretion.

The nephridia empty into mantle cavity.

Annelids (Earthworm):

 

The excretory system consists of paired nephridia or coiled tubules in each segment.

Nephridia have 2 openings: one is a ciliated funnel that collects coelomic fluid and the other is an exit in the body wall.

Between the 2 openings is a convoluted region where waste material is removed from the blood vessels about the nephridium.

Arthropods (Insect):

 

Terrestrial forms have an excretory system that conserves water.

The malphigian tubules collect wastes from the blood and reabsorb water so that only dry material is excreted.

Marine crustaceans such as gry fish have excretory system consisting of a pair of green glands lying in the head region anterior to the oesophagus.

Each organ possesses a glandular region for waste removal, a bladder and a duct that opens ventrally at base of the antennae.

Echinoderms (Starfish):

 

Star fish has no excretory organs.

Chordates:

 

The kidneys are important excretory and water regulating organs that conserve or rid the body of water as appropriate.

Fishes:

 

The kidney filters liquid waste materials from the body, these wastes are then passed out of the body.

The kidney is also extremely important in regulating water and salt concentration within the fish’s body, allowing certain fish species to exist in freshwater or salt water and in some cases both.

The vent is the external opening to digestive, urinary and reproductive tracts. In most fish it is immediately in front of anal fin.

Ammonia is formed immediately after the amino group is removed from protein. This process requires very little energy. Ammonia is highly soluble in water but very toxic.

Aquatic animals such as bony fishes, aquatic invertebrates and amphibians excrete ammonia because it is easily eliminated in the water.

Amphibians:

 

Liquid wastes from kidney travel by way of ureters to the urinary bladder.

Solid wastes from the large intestine pass into the cloaca.

Both liquid and solid waste materials leave the body by way of the cloaca and the cloacal vent.

Terrestrial amphibians and mammals excrete nitrogenous wastes in the form of urea because it is less toxic than ammonia and can be moderately concentrated to conserve water.

Urea is produced in the liver by a process that requires more energy to produce than ammonia does.

Reptiles:

 

The metanephric kidneys of reptiles are lobulated. One or more renal arteries can be presented to receive blood from the renal portal system.

The nitrogenous wastes of reptiles are in form of ammonia, urea, uric acid or a combination of these.

Crocodilians, snakes and some lizards do not have a urinary bladder.

In chelonians and lizards with a bladder is connected to the cloaca by a short urethra.

Urine passes into the cloaca and then into the urinary bladder if present or into distal colon where water resorption occurs.

The cloaca typically consists of 3 chambers –

The coprodeum is the most cranial and receives faecal material and urinary wastes.

The uroderm is the middle section and receives genital secretions and urinary wastes from the urogenital ducts.

The caudal proctoderm acts as a reservoir for faecal and urinary wastes before they are excreted. This also is the location of the openings of the musk glands.

Birds: 

 

Insects, reptiles, birds and some dogs (Dalmatians) excrete uric acid.

Reptiles and birds eliminate uric acid with their faces.

The white material seen in bird droppings is uric acid.

It is not very toxic and is not very soluble in water.

Excretion of wastes in the form of uric acid conserves water because ti can be produced in a concentrated form due to its low toxicity.

Because it is relatively insoluble and nontoxic, it can accumulate in eggs without damaging the embryo.

The synthesis of uric acid requires more energy than urea synthesis.

There is no urinary bladder in birds.

Mammals:

 

In mammals, 2 major excretory processes are the formation of urine in the kidneys and the formation of faces in the intestine.

The waste products are eliminated by urination and defecation respectively.

While urine and faces are both water material of body processes they are in completely different categories.

Urine is a waste product of the urinary system process while faces are waste products of digestive system.

Faces may contain harmful materials such as bacteria, viruses and parasitic worms.

Urine on the other hand, contains excess water, salt and protein waste in the form of urea and seldom carries any pathogens.

Primates:

The kidney is a major excretory organ of primates and other vertebrates.

The principle responsibility of the organ is to separate urea, toxins and other types of waste from the blood, while water; salt and electrolytes are maintained at an appropriate level.

Due to this important role, the kidney is also involved in blood pressure and acid base regulation in the body.

Nephrons are the basic filtering units of the kidney, more than a million of them being present in a normal adult human kidney.

Working together, the nephrons are able to filter blood at an impressive rate, processing the entire 5 quarter water content of human circulatory system about every 45 minutes.

Only a minute portion of the material passing through the kidneys is actually excreted, however, the vast majority being reabsorbed by the nephrons.